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It uses very old language that is more confusing today than helpful, including references to `SubstNt` that no longer exists. The comment above `TokenStream` is better, and suffices for a basic understanding of these types.
The rustc_ast crate contains those things concerned purely with syntax
– that is, the AST ("abstract syntax tree"), along with some definitions for tokens and token streams, data structures/traits for mutating ASTs, and shared definitions for other AST-related parts of the compiler (like the lexer and macro-expansion).
For more information about how these things work in rustc, see the rustc dev guide: